The Dutch-led joint investigation team (JIT) named the men as Igor Girkin, Sergei Dubinsky and Oleg Pulatov from Russia, as well as Ukrainian Leonid Kharchenko.
The JIT, which is seeking to try the suspects under Dutch law, had previously said it had a "long list" of persons of interest and appealed again for witnesses as the investigation continues.
Who are the suspects?
The most prominent of the four is Igor Girkin (also known as Strelkov), who prosecutors say is a former colonel in Russia's FSB intelligence service. He was given the minister of defence title in the rebel-held eastern Ukrainian city of Donetsk.
He is believed to be the highest military officer in the area who was in direct contact with the Russian Federation. In a statement Mr Girkin said: "I can only say that militia did not shoot down the Boeing."
Image copyrightREUTERSImage captionAn investigator inspects the wreckage of Malaysia Airlines Flight MH17
The others charged are:
Sergei Dubinsky (known as Khmury), who prosecutors say was employed by Russia's GRU military intelligence agency, was a deputy of Mr Girkin and was in regular contact with Russia
Oleg Pulatov, known as Giurza, who the JIT says was a former soldier of GRU special forces and deputy head of the intelligence service in Donetsk
Ukrainian national Leonid Kharchenko, who has no military background but led a combat unit as a commander in Eastern Ukraine, according to prosecutors
"The JIT suspects Girkin, Dubinsky, Pulatov and Kharchenko of having played a significant role in the death of 298 innocent civilians," said prosecutor Fred Westerbeke.
"The means of evidence these suspicions are based on will be elaborated on during the criminal proceedings, then the judge will pass sentence. The suspects are innocent until the contrary is proven in court."
He added that investigators had "evidence showing that Russia provided the missile launcher".
The passenger jet left Amsterdam's Schiphol Airport at 10:15 GMT on 17 July 2014 and was due to arrive at Kuala Lumpur in Malaysia the following day.
A few hours after take-off, the plane lost contact with air traffic control about 50km (30 miles) from the Russia-Ukraine border.
At the time, an armed conflict was raging on the ground in eastern Ukraine between Russian-backed separatists and Ukrainian government forces, and several government military aircraft had been downed in the previous weeks, while air strikes were carried out on rebel-held areas.
The plane crashed in the Donetsk area, in territory controlled by separatists. Parts of the wreckage were found distributed over an area of about 50sq km (19 square miles).
In October 2015, the Dutch Safety Board concluded the plane had been hit by a Buk missile, causing it to break apart in mid-air.
Media captionAn animated video from the Dutch Safety Board shows the damage to the plane and how it was caused
Russia responded by denying any of its anti-aircraft missile systems had ever crossed the Ukrainian border. Its foreign ministry has accused the JIT investigation of being "biased and politically motivated".
UK Foreign Secretary Jeremy Hunt said Russia "must co-operate fully with the prosecution and provide any assistance it requests" in accordance with UN Security Council resolution 2166, which was passed in response to the downing of MH17.
"The international community stands together against the impunity of those responsible for the despicable murder of 298 innocent people," he added.
Nato chief Jens Stoltenberg said the bringing of charges marked an "important milestone in the efforts to uncover the full truth and ensure that justice is done".
Who were the victims?
A total of 283 passengers, including 80 children, and 15 crew members were killed on the flight.
Image copyrightVARIOUSImage captionThose killed were from across the world
The dead included 193 Dutch nationals, 43 Malaysians, 27 Australians, 12 Indonesians, 10 Britons, four Belgians, four Germans, three Philippine nationals, one New Zealander and one Canadian.
Image copyrightREUTERSImage caption"It's a start" - Silene Fredriksz, mother of one of the victims
Speaking ahead of Wednesday's announcement, Silene Fredriksz, who lost her son Bryce, said that in the five years since the downing of MH17, some relatives had died not knowing the truth.
"We all get older... I hope that I will know the truth before I close my eyes," she said.
Delays and difficulties in the recovery of victims' remains prompted widespread international criticism.
Keep these in mind as you contemplate the direction of the American government over the past 50 years and especially since the Obama election.
The Goals of Communism
(as read into the congressional record January 10, 1963, from "The Naked Communist" by Cleon Skousen)
1. U.S. acceptance of coexistence as the only alternative to atomic war.
2. U.S. willingness to capitulate in preference to engaging in atomic war.
3. Develop the illusion that total disarmament of the United States would be a demonstration of moral strength.
4. Permit free trade between all nations regardless of Communist affiliation and regardless of whether or not items could be used for war.
5. Extension of long-term loans to Russia and Soviet satellites.
6. Provide American aid to all nations regardless of Communist domination.
7. Grant recognition of Red China. Admission of Red China to the U.N.
8. Set up East and West Germany as separate states in spite of Khrushchev's promise in 1955 to settle the German question by free elections under supervision of the U.N.
9. Prolong the conferences to ban atomic tests because the United States has agreed to suspend tests as long as negotiations are in progress.
10. Allow all Soviet satellites individual representation in the U.N.
11. Promote the U.N. as the only hope for mankind. If its charter is rewritten, demand that it be set up as a one-world government with its own independent armed forces. (Some Communist leaders believe the world can be taken over as easily by the U.N. as by Moscow. Sometimes these two centers compete with each other as they are now doing in the Congo.)
12. Resist any attempt to outlaw the Communist Party.
13. Do away with all loyalty oaths.
14. Continue giving Russia access to the U.S. Patent Office.
15. Capture one or both of the political parties in the United States.
16. Use technical decisions of the courts to weaken basic American institutions by claiming their activities violate civil rights.
17. Get control of the schools. Use them as transmission belts for socialism and current Communist propaganda. Soften the curriculum. Get control of teachers' associations. Put the party line in textbooks.
18. Gain control of all student newspapers.
19. Use student riots to foment public protests against programs or organizations which are under Communist attack.
20. Infiltrate the press. Get control of book-review assignments, editorial writing, policymaking positions.
21. Gain control of key positions in radio, TV, and motion pictures.
22. Continue discrediting American culture by degrading all forms of artistic expression. An American Communist cell was told to "eliminate all good sculpture from parks and buildings, substitute shapeless, awkward and meaningless forms."
23. Control art critics and directors of art museums. "Our plan is to promote ugliness, repulsive, meaningless art."
24. Eliminate all laws governing obscenity by calling them "censorship" and a violation of free speech and free press.
25. Break down cultural standards of morality by promoting pornography and obscenity in books, magazines, motion pictures, radio, and TV.
26. Present homosexuality, degeneracy and promiscuity as "normal, natural, healthy."
27. Infiltrate the churches and replace revealed religion with "social" religion. Discredit the Bible and emphasize the need for intellectual maturity which does not need a "religious crutch."
28. Eliminate prayer or any phase of religious expression in the schools on the ground that it violates the principle of "separation of church and state."
29. Discredit the American Constitution by calling it inadequate, old-fashioned, out of step with modern needs, a hindrance to cooperation between nations on a worldwide basis.
30. Discredit the American Founding Fathers. Present them as selfish aristocrats who had no concern for the "common man."
31. Belittle all forms of American culture and discourage the teaching of American history on the ground that it was only a minor part of the "big picture." Give more emphasis to Russian history since the Communists took over.
32. Support any socialist movement to give centralized control over any part of the culture--education, social agencies, welfare programs, mental health clinics, etc.
33. Eliminate all laws or procedures which interfere with the operation of the Communist apparatus.
34. Eliminate the House Committee on Un-American Activities.
35. Discredit and eventually dismantle the FBI.
36. Infiltrate and gain control of more unions.
37. Infiltrate and gain control of big business.
38. Transfer some of the powers of arrest from the police to social agencies. Treat all behavioral problems as psychiatric disorders which no one but psychiatrists can understand.
39. Dominate the psychiatric profession and use mental health laws as a means of gaining coercive control over those who oppose Communist goals.
40. Discredit the family as an institution. Encourage promiscuity and easy divorce.
41. Emphasize the need to raise children away from the negative influence of parents. Attribute prejudices, mental blocks and retarding of children to suppressive influence of parents.
42. Create the impression that violence and insurrection are legitimate aspects of the American tradition; that students and special-interest groups should rise up and use united force to solve economic, political or social problems.
43. Overthrow all colonial governments before native populations are ready for self-government.
44. Internationalize the Panama Canal.
45. Repeal the Connally reservation so the United States cannot prevent the World Court from seizing jurisdiction over nations and individuals alike.
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